We definitely have an identify problem here.
You say you have a 2N5485 in Q1. That is a FET that should be used in Q2 through Q6. Q1 is supposed to be a NPN transistor BC550.
You say you are using MPSA18 in Q2 through Q6. Those are NPN transistors. You need 2N5485 in Q2 through Q6.
Your voltages are all wrong. You need to have about 9V on the drains of Q2, Q4 and Q6.
Correct the transistors and remeasure. Be sure you are measuring correctly. Consult our guides for more information.